Duane Keith Davis, charged with murder in Tupac Shakur's 1996 death, pleads not guilty in Las Vegas

Duane Keith "Keffe D" Davis, a former Southern California street gang leader, pleaded not guilty Thursday to murder in the 1996 killing of rap music icon Tupac Shakur in Las Vegas — a charge prompted by his own descriptions in recent years about orchestrating the deadly drive-by shooting.

Davis is the only person still alive who was in the vehicle from which shots were fired and the only person ever charged with a crime in the case.